Cyclosia eucharia Jordan
Cyclosia eucharia Jordan, 1907, Gross-Schmett. Erde, 10: 20.
Diagnosis
As in inclusus, both this species and hecabe Jordan are markedly sexually dimorphic. C. eucharia males have the wings generally dark brown, though becoming greyer basally. The forewings have two well separated pale brown patches submarginally. The more distal of these patches is represented in the female by a deep indentation to the black border that is typical of Eurema species. C. hecabe males have blue hindwings and an oblique, almost continuous, pale greyish subapical band to the blackish forewing; the females have two closely associated pale spots within the black border of the forewing that resemble the more distal part of the pale band in the male.
Taxonomic note
Jordan (1907) indicated some confusion in his original text over the assignation of males of females, and included eucharia females with the only hecabe male in his description of the latter, initially emphasising the female characteristics in his text. However, he referred to a series of one male from G. Marapok and two females from G. Marapok and Penang effectively as a syntypic series. He subsequently labelled the males illustrated as holotypes of his two species, but did not label any females with type status. Therefore, the male with blue hindwings from G. Marapok is here formally designated as LECTOTYPE of hecabe and the male from Penang, with dark brown forewings with two well separated paler brown patches, is here formally designated as the LECTOTYPE of eucharia. On this basis, the taxa are definitely not conspecific as stated by Endo & Kishida (1999), though these authors correctly associated males and females of the species here treated as eucharia; both sexes of eucharia have been recorded from both Borneo and Penang. The two species were recognised as distinct by Yen et al. (2005), though they (fig 22Z-2) illustrated a female of eucharia as hecabe. The pairing of sexes here is based on concordance in shape and position of the pale forewing markings of the males nearest the distal margin with: the indentation to the inner border of the black margin in eucharia; the pair of pale spots enclosed within the black margin in hecabe.
Geographical range
Borneo, Peninsular Malaysia (Penang).
Habitat preference
The species is recorded from G. Marapok and G. Kinabalu in Sabah without precise data. There is also a female taken from undergrowth in secondary forest on the trail to G. Dulit in Sarawak, though no altitude is given, and another (in FRC, Sepilok) from about 500m at Moyog (Kipandi) near Kota Kinabalu in Sabah.
Genitalia:
