Cyclosia Hübner

Type species: panthona Stoll, China, Indian Subregion.

Synonyms:

• Epyrgis Herrich-Schäffer (type species midama)
• Callamesia Butler (type species midama Herrich-Schäffer)
• Didina Walker (type species thecloides Walker)
• Isbarta Walker (type species glauca Walker = pieridoides Herrich-Schäffer)
• Klaboana Moore (type species macularia Guérin-Méneville).

This genus includes a diversity of facies types, with sexual dimorphism that ranges from minor to extreme and may involve one (usually the female) or both sexes in mimicry of diverse other lepidoptera groups; such as Pieridae, Danainae, and day-flying Arctiinae. Two features in particular facilitate recognition of the genus but are not absolutely exclusive to it: presence of a series of small cross-veins between the costa and subcosta of the forewing (also in Eucormopsis and a subclade of Clade 18 (see p. 78)); the general form of the male genitalia, particularly a spur from each side of the tegumen. The genitalia are elongate compared to most Chalcosiinae, the valves much longer than they are deep. The uncus is entire, narrow, tapering, slightly hooked. The saccus is usually well developed. The aedeagus is slender, slightly curved and as long as, to very much longer than, the valves; this is extreme in C. inclusus Walker where the aedeagus is flexed back on itself and probably is in total much longer than the abdomen. The female ovipositor is strongly extensile.

Robinson et al. (2001) recorded the type species as feeding on Aporusa, Baccaurea (Euphorbiaceae) and Hydnocarpus (Flacourtiaceae). They listed the same euphorb genera for C. papilionaris Drury (mainland Asia) but also Dipterocarpus (Dipterocarpaceae), Castanopsis, Quercus (Fagaceae), Solanum (Solanaceae) and also unidentified Rutaceae. Two Bornean species (see below) have also been recorded from Euphorbiaceae.