Eutoeini

The male genitalia have the valve almost completely divided, the upper arm being setose: the costal margin of typical ennomines. Unlike the Macariini, the costa is rarely delimited within it by a longitudinal ridge, and the transtilla is weak. The ventral arm is as long as the costal one, slender, terminating acutely in a gentle curve. The uncus is setose, a short, blunt apical hook arising dorsally in most genera from rounded to square lateral ‘shoulders’. The gnathus is narrow, straplike, or vestigial. The aedeagus vesica usually contains one or two large, sometimes elaborate, cornuti, rarely a cluster of smaller ones (e.g. Eutoea Walker).

The male abdomen has a transverse setal comb on the third sternite and occasionally (particularly in Eutoea, Zeheba Moore and Luxiaria Walker) a central apodeme-like structure on the anterior margin of sternite 7. The male antennae are bipectinate to ciliate, the latter state occurring throughout the three genera just mentioned. These genera share other features such as a lobe-like structure separating the two arms of the valve, a flange or flanges subapically on the ventral arm of the valve, and boss of the signum in the bursa copulatrix of the female. They probably represent a natural group within the tribe.

No features of the female genitalia appear to be definitive, though the trio of genera just mentioned has lost the signum.

As well as the genera discussed below, the tribe includes the Australasian genera Nadagarodes Warren (possible sister-group to ProbithiaWarren, as reviewed by Holloway (1984b)) and Orthotmeta Warren (four species in New Guinea) and the African genera Pigiopsis Warren, IdiotephraWarren (distinct from Luxiaria) and Anoectomychus Prout.

No features of early stages appear to be of tribal significance. Sevastopulo (1947) gave a brief account of the larva of the type species of Bithiodes, obliquata Moore. The pupa is subterranean, and the host-plant is Melastoma. One of the two other records for the tribe (See Calletaera postvittata Walker) is also from the Melastomataceae.