Holcocerus Staudinger

Schoorl (1990: 54‑55) treated this genus as distinct, but his analysis placed it as sister‑group to Cossus. The male and female antennae are prismatic in Holcocerus (referred to as filiform in Part 1) but each flagellomere is unilobed in Cossus. In this, both genera show a reduction, treated by Schoorl as synapomorphy, from the plesiomorphic bipectinate condition. There is also a synapomorphy in the basisternum of the stenopleural region of the mesothorax: this sclerite is shortened and rounded posteriorly. Schoorl placed two Bornean species in Holcocerus, and the antennae of chloratoides Holloway indicate it may also belong in this genus, comb. n.

H. chloratoides has been recorded from Peninsular Malaysia (FRIM colln). Holcocerus as recognised by Schoorl extends to Eurasia and Africa, but is not found further east than Sundaland in the Oriental region.