Cleoda Tremewan

Type species: syntomoides Walker, Peninsular Malaysia.

Synonym: Doclea Walker (type species syntomoides), praeocc.

Specimens of both sexes attributed to this genus have long, narrow wings, with radial, medial and anterior cubital veins arising directly from the cell in the forewing. CuA2 is convergent on CuP at the margin on the hindwing. The male antennae are strongly bipectinate, black, the rami somewhat paddle-like. The male abdomen is pilose, with a prominent black apical tuft. The colouring is generally dark brown to black, with greenish tints on the forewings.

The female is larger with more rounded wings apices and has more extensive areas of pale coloration, that of the type species being extensively dull ochreous yellow on both surfaces of the hindwing, with a narrow brown margin. Yen et al. (2005) discussed in some detail whether the female indicated to be the type of syntomoides in OUMNH is related to males in BMNH that accord with Walker’s original description of a specimen of that sex from Malakka with a greenish forewing. However, his description of the hindwing matches that of the female closely, rather than the mostly dark hindwings of those males, so it is probable that his attribution of sex was an error. A male from Borneo in the collection of Thomas Witt (Museum Witt, Munich) has ochreous hindwings as in the Oxford female, and this may offer a more likely marriage. Jordan (1907) indicated that a male of syntomoides from Java in OUMNH was tinged yellowish where those from Peninsular Malaysia are reddish.

The male abdomen has the sclerites of the eighth segment reduced, and there is a pair of elongated scaled coremata between this segment and the genitalia. The genitalia have the uncus divided into a pair of rectangular plates. The tegumen is much longer than the very shallow but broad vinculum as illustrated. The valves are simple, ovate, entire, with a zone of scaling over the interior of the apical part. The aedeagus is slender and slightly sinuous, to the apex.

The female abdomen has not been examined, but Yen et al. (2005: 272) stated that the ovipositor is extended.

The genus consists of the type species and a new one from Borneo described below. A further new Bornean species with atypical male genitalia is also tentatively included.