Caelestomorpha velutina Walker comb. n.
Narosa velutina Walker, 1862, J. Linn. Soc. Lond. (Zool.) 6: 171.
Diagnosis
The species is smaller than concinna (11-12mm). The forewings are squarer, less produced apically, with the darker orange zone subbasally and the postmedial bar less definitely divided by a paler fascia or zone. The main difference from concinna is in the male genitalia where the gnathus is less deeply bifid, the juxta and associated structures smaller, less heavily spiralled and apically more finely spined. The genitalia of two specimens are illustrated to show the variations.
Habitat Preference
The species is known mainly from lowland localities in rainforest, the highest record being from Kiau at 1000m on G. Kinabalu.
Taxonomic Notes
The Sulawesi species attributed to Narosa by Holloway (1987a) are related to the Sundanian species concinna Swinhoe and velutina Walker, both occurring in Borneo. The group also includes an undescribed species from Waigeu and pectinata Hering from New Guinea. A preliminary phylogenetic analysis (Holloway, 1987a) indicated that the Sundanian pair was sister to a grouping of all the more easterly species. This sister pair of clades in turn related to the Himalayan endodonta Hampson. Solovyev & Witt (2009) described a new genus, Caelestomorpha Solovyev & Witt, with type species albiceris Solovyev & Witt from Thailand, Cambodia and Vietnam, but included endodonta as a second species. Despite the strong and asymmetric development of the saccular processes of the valves in taxa from Sundaland eastwards, the valve shape and structure of the uncus and gnathus are as in the generic definition of Solovyev & Witt, and so all named taxa from this more eastern complex are transferred to Caelestomorpha: *C. concinna Swinhoe comb. n.; C. velutinacomb. n.**; C. pectinata Hering comb. n. None of the Sulawesi species has been described, but the phylogenetic analysis of Holloway (1987) was discussed further, with more ecological detail, in the context of speciation on Sulawesi by Holloway (1991).
Genitalia:
